Research articles

  1. R. Azzarelli, C. Hurley, M. K. Sznurkowska, S. Rulands, L. Hardwick, I. Gamper, F. Ali, L. McCracken, C. Hindley, F. McDuff, S. Nestorowa, R. Kemp, K. Jones, B. Göttgens, M. Huch, G. Evan, B. D. Simons, D. Winton, A. Philpott, 
    Multi-site Neurogenin3 phosphorylation controls pancreatic endocrine differentiation,
    Developmental Cell 41(3), 274-286.e5 (2017)
  2. M. Aragona*, S. Dekoninck*, S. Rulands, S. Lenglez, G. Mascré, B. D. Simons, C. Blanpain, 
    Uncoupling stem cell proliferation, differentiation and migration during wound healing,
    Nature Communications, 14684 (2017)
  3. S. Rulands, B. D. Simons,
    Tracing cellular dynamics in tissue development, maintenance and disease,
    Current Opinion in Cell Biology 43, 38–45 (2016)
  4. A. Wuidart*, M. Ousset*, S. Rulands, B. D. Simons, A. Van Keymeulen, C. Blanpain,
    Quantitative lineage tracing strategies to resolve multipotency in tissue specific stem cells,
    Genes and Development 30, 1261-1277 (2016)
  5. Y. Wan, A. D. Almeida, S. Rulands, N. Chalour, L. Muresan, Y. Wu, B. D. Simons, J. He, W. A. Harris,
    The ciliary marginal zone of the zebrafish retina: clonal and time-lapse analysis of a continuously growing tissue,
    Development, 0950-1991 (2016)
  6. S. Chabab*, F. Lescroart*, S. Rulands*, N. Mathiah, B. D. Simons, C. Blanpain
    Uncovering the number and clonal dynamics of Mesp1 progenitors during heart morphogenesis,
    Cell Reports 14(1), 1-10 (2016)
  7. H. Boije, S. Rulands, S. Dudczig, B. D. Simons, W. A. Harris
    The independent probabilistic firing of transcription factors: A paradigm for clonal variability in the zebrafish retina,
    Developmental Cell, 34(5), 532–543 (2015)
  8. J. K. Watson, S. Rulands, A. C. Wilkinson, A. Wuidart, M. Ousset, A. Van Keymeulen, B. Göttgens, C. Blanpain, B. D. Simons, E. L. Rawlins
    Clonal dynamics reveal two distinct populations of basal cells in slow turnover airway epithelium,
    Cell Reports, 12(1), 90-101 (2015).
    Gurdon Institute news item
  9. F. Lescroart, S. Chabab, X. Lin, S. Rulands, C. Paulissen, A. Rodolosse, H. Auer, Y. Achouri, C. Dubois, A. Bondue, B. D. Simons, C. Blanpain,
    Early lineage restriction in temporally distinct populations of Mesp1 progenitors during mammalian heart development,
    Nature Cell Biology 16, 829-840 (2014).
  10. S. Rulands, D. Jahn, E. Frey,
    Specialization and bet hedging in heterogeneous populations,
    Physical Review Letters 113, 108102 (2014). [arxiv:1408.4912]
  11. M. Reiter*, S. Rulands*, E. Frey,
    Range expansion of heterogeneous populations,
    Physical Review Letters 112, 148103 (2014). [arxiv:1403.6333]
    LMU press release (German), NIM press release
  12. S. Rulands, A. Zielinski, E. Frey,
    Global attractors and extinction dynamics of cyclically competing species,
    Physical Review E 87, 052710 (2013). [arxiv:1305.0119]
  13. S. Rulands, B. Klünder, E. Frey,
    Stability of localized wave fronts in bistable systems,
    Physical Review Letters 110, 038102 (2013). [arxiv:1212.1036]
    LMU press release
  14. S. Rulands, T. Reichenbach, Erwin Frey,
    Three-fold way to extinction in populations of cyclically competing species,
    Journal of Statistical Mechanics L01003 (2011). [arXiv:1005.5704]

* denotes equal contributions.


  • S. Rulands
    Heterogeneity and spatial correlations in stochastic many-particle systems.
    From embryogenesis to evolution.
    PhD Thesis, Ludwig-Maximilians-University Munich (2013)
    Advisor: Prof. Erwin Frey
  • S. Rulands
    Statistics of rare events in reaction-diffusion systems
    Diploma Thesis, Ludwig-Maximilians-University Munich (2009)
    Advisor: Prof. Erwin Frey

Lecture notes

  • Tutorial on advanced pattern formation - Amplitude equations and wave fronts,
    Theory of Living Matter group, Cambridge, September 2014 
    Lecture notes (PDF)